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The film is set in the 1750s and involves the Jesuit Reductions, a programme by which the European Catholic Church sought to Christianise and "civilise" the indigenous native populations of South America. It tells the story of a Spanish Jesuit priest, Father Gabriel (Jeremy Irons), who enters the South American jungle to build a mission and convert a community of Guaraní Indians to Christianity, and of the eventual destruction of the missions by the secular Spanish and Portuguese colonial governments.

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